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Ph.D. Wildlife Biology, University of Montana, 2010, Dissertation, ‘Escaping the extinction vortex: identifying factors affecting population performance and recovery in endangered Sierra Nevada bighorn sheep’; M.Sc. Wildlife Ecology, University of Arizona, 2003, Thesis, ‘Antler breakage in tule elk: nutritional causes and behavioral consequences’; B.Sc. Biology – Ecology, Behavior and Evolution, University of California San Diego, 1999.

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Wildlife Researcher, Colorado Parks and Wildlife, 2010 to present; Post-Doctoral Researcher, USGS Montana Cooperative Wildlife Research Unit, University of Montana; Wildlife Biologist, California Department of Fish and Game.

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Black bear use of urban environments - testing management strategies to reduce bear-human conflicts and assessing the role of human development on bear behavior and population dynamics

Finding solutions for reducing elk and deer damage to high-value agricultural resources

Examining the relative influence of habitat, weather, land-use, and predation on adult female elk survival and cause-specific mortality across the western U.S.

Identifying the demographic, habitat, predation, and genetic factors affecting population performance and recovery in federally endangered Sierra Nevada bighorn sheep

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population modeling, resource selection, conservation genetics, predator-prey relationships, Bayesian applications in ecology, wildlife-human conflicts, endangered species conservation, large mammal ecology and management

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University of Montana Wildlife Biology Program Graduate Student Research Award, 2010

P.E.O. Scholar Award, 2009

Canon National Park Science Scholarship Dissertation Award, 2007

National Fish and Wildlife Foundation/Budweiser Conservation Scholarship, 2006

University of Arizona, School of Renewable Natural Resources, Best Thesis Award, 2004

University of California, Graduate Student Research Award, 2003

Seegmiller Foundation Scholarship, University of Arizona, 2002

Hansen-Welles Scholarship, Desert Bighorn Council, 2002

Member of The Wildlife Society, Ecological Society of America, and Society for Conservation Biology

Reviewer for Journal of Wildlife Management, Conservation Biology, Behavioral Ecology, Journal of Mammalogy, and Conservation Genetics

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MILLS, L.S., AND H.E. JOHNSON. Wildlife Population Dynamics. Book chapter in textbook “Wildlife management: contemporary principles and practices.” John Hopkins University Press. In Press.

JOHNSON, H.E., M. HEBBLEWHITE, T.R. STEPHENSON, AND D. GERMAN. Evaluating apparent competition in limiting the recovery of an endangered ungulate.  Oecologica: In Press.

BRODIE, J., H. JOHNSON, M. MITCHELL, P. ZAGER, K. PROFFITT, M. HEBBLEWHITE, M. KAUFFMAN, B. JOHNSON, J. BISSONETTE, C. BISHOP, J. GUDE, K. HERSEY, M. HURLEY, P. LUKACS, S. McCORQUODALE, E. McINTIRE, J. NOWAK, D. SMITH, AND P.J. WHITE. Relative influence of human harvest, carnivores, and weather on adult female elk survival across western North America. Journal of Applied Ecology: In Revision.

JOHNSON, H.E., L.S. MILLS, J.D. WEHAUSEN, T.R. STEPHENSON, AND G. LUIKART. 2011. Translating effects of inbreeding depression on component vital rates to overall population growth in endangered bighorn sheep. Conservation Biology 25:1240-1249.

CAHN, M.L., M.M. CONNER, O.J. SCHMITZ, M.W. MILLER, T.R. STEPHENSON, H.E. JOHNSON, AND J.D. WEHAUSEN. 2011. Disease, population viability, and recovery of endangered Sierra Nevada bighorn sheep. Journal of Wildlife Management 75:1753-1766.

JOHNSON, H.E., L.S. MILLS, J. WEHAUSEN, AND T.R. STEPHENSON. 2010. Population-specific vital rate contributions influence management of an endangered ungulate. Ecological Applications 20:1753-1765.

JOHNSON, H.E., L.S. MILLS, J. WEHAUSEN, AND T.R. STEPHENSON. 2010. Combining ground count, telemetry, and mark-resight data to infer population dynamics in an endangered species. Journal of Applied Ecology 47:1083-1093.

BLEICH, V.C., H.E. JOHNSON, S.A. HOLL, L. KONDE, S.G. TORRES, AND P.R. KRAUSMAN. 2008. Fire history in a chaparral ecosystem: implications for conservation of a native ungulate. Rangeland Ecology and Management 61:571-579.

JOHNSON, H.E., V.C. BLEICH, AND P.R. KRAUSMAN. 2007. Mineral deficiencies in tule elk, Owens Valley, California. Journal of Wildlife Diseases 43:61-74.

JOHNSON, H.E., V.C. BLEICH, AND P.R. KRAUSMAN, AND J.L. Koprowski. 2007. Effects of antler breakage on mating behavior in male tule elk. European Journal of Wildlife Research 53:9-15.
 
CAIN, J.W., H.E. JOHNSON, AND P.R. KRAUSMAN. 2005. Wildfire and desert bighorn sheep habitat. Southwestern Naturalist 50:506-513.
 
JOHNSON, H.E., V.C. BLEICH, AND P.R. KRAUSMAN. 2005. Antler breakage in tule elk, Owens Valley, California. Journal of Wildlife Management 69:1747-1752.